While reading a magazine on an airplane to Acapulco, Frasier notices the model seated next to him is the model on the magazine. The two embark on an affair only under the conditions Frasier remains secretive about their relationship. When Frasier returns to Seattle, no one believes him about the Model and think she’s possibly a figment of his imagination.
With their father’s 65th birthday approaching, Frasier and Niles compete to buy Martin the best gift. While Frasier settles on a new television, Niles opts to buy him his old police horse.
In the season of Halloween, Niles decides to throw an elaborate ‘literary’ themed costume party. With the entire crew in attendance, Roz makes a shocking announcement to Frasier that she may be pregnant.
Now that Roz has revealed her pregnancy, she tells Frasier who the father is, a man much younger than she is. As Frasier urges her to speak with the father about the pregnancy, Roz’s dilemma grows as she makes a decision to keep or not keep it.
Upon the 1000th episode of Frasier’s ‘call-in’ radio program, Mayor Norman Rice designates ‘Frasier Crane Day’. Celebrating with a gala at the Seattle Space Needle, everything is in order minus one key ingredient, Frasier.
Frasier sets sail on a ‘celebrity’ cruise line with Niles and Roz in tow. Not only do they discover that luxury is only in its name, but also Maris, Niles estranged wife, is booked on the same cruise.
Frasier falls for a high-profile criminal defense attorney, Samantha. He finds that his usually controlling self is at the mercy of his dominant lover.
When Samantha, Frasier’s romantic interest from the previous episode, breaks off their relationship, she swears to him that it is ‘her’ not ‘him’. Frasier is unconvinced otherwise and becomes obsessed that the break-up is entirely his fault.
Frasier, Niles, Daphne and Roz visit the office of their favorite masseuse who massages out the tensions from their holiday season. Each rehashes their stressful events including Martin partaking in a Christmas pageant and Niles getting trapped in an elevator.
Missing the welcoming atmosphere of Cheers, Frasier comes across a similar Seattle pub called the Fox and Whistle. Enjoying the chummy atmosphere with the regulars, Daphne becomes agitated as this bar happens to be her stomping ground.
When Daphne discovers an engagement ring in Martin’s bedroom, she fills in Frasier and Niles on the news. Not just disturbed by the idea of Sherry as their stepmother, Frasier and Niles also find out that Sherry has quite a list of previous marriages.
It is contract negotiation time at the station and Frasier decides to dismiss his agent Bebe, who is pushing for a bigger salary so she receives a bigger commission. Instead, he teams up with Ben, an honest rep, who wants the Seattle Zoo to name a bird ‘Frasier Crane’.
Niles and his estranged wife Maris begin marriage counseling with a new counselor, Dr. Schenkman. When the doctor suggests that Niles becomes more spontaneous in the bedroom, Niles is unprepared for his discovery.
When Frasier, Niles and Martin take a weekend getaway to a ski lodge, they receive the company of Daphne and her attractive friend Annie. A snow pile of mix-ups leads to a disastrous weekend filled with misunderstandings.
Lilith comes to Seattle to inform Frasier that her husband left her and she winds up seducing Niles, changing Frasier and Lilith's relationship forever.
Since Frasier’s advice to his cousin Nikos to give up a career in medicine to pursue juggling, Frasier and Nikos’ mother, Zora, have not spoken. Now that Nikos is about to get married, Zora shows up at Frasier’s apartment looking to make mends or so he thinks.
A new ‘call-in’ radio host at the station leaves a mad taste in Frasier’s mouth. Dr. Crane fears that Dr. Clint Webber, slick and handsome, will become a permanent fixture at KACL. Meanwhile, Niles must reconcile between his father and a gourmet pet store who refuses to let Martin back in.
Bulldog stops a gunman from holding up a coffee shop when he accidently throws coffee on him. To the patrons, Bulldog is a hero and receives much acclaim for his good deed. Frasier however knows the truth and the attention Bulldog receives is killing him.
When Frasier embarks on a heated romance with Caitlin, an earthy artist, he tries to convince the others that his attraction for her is on her talent not just physical. The idea is hard to believe though since the two can’t keep their hands off one another.
Now that the marriage between Niles and Maris is completely over, he decides to ask Daphne out to dinner. Too chicken to ask, he pretends to have another woman over for dinner but his plan almost backfires as Daphne shows up offering to cook the meal for them.
Roz’s baby is coming soon and she begins to worry about its appearance as the baby’s father, Rick, has a family of large noses. Meanwhile, Frasier gives Daphne a pair of sapphire earrings but worries since he did not tell her they are fake.
To overcome the frustration of the single life, Frasier and Niles throw a singles party. With the very pregnant Roz in attendance, the party hits a high point when the mother-to-be goes into labor.
On the night that Frasier is to meet up with a very attractive date, he is torn between his obligations to attend a Safari Club initiation and a birthday party. And not immune to women troubles, Niles falls head over heels for a woman too sophisticated for even his taste.
Frustrated with the new station manager’s request to perform live commercials for various products, Frasier unites his co-workers against the policy. Unfortunately his protest does not go as planned and his job may possibly be on the outs.
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